Authors(s)	Dowler, R. C., and J. W. Bickham.
Year	1982
Title	Chromosomal relationships of the tortoises (family Testudinidae). Genetica 58: 189-197.

• Geochelone pardalis
• Gopherus agassizii
• Gopherus berlandieri
• Gopherus flavomarginatus
• Gopherus polyphemus
• Indotestudo elongata
• Malacochersus tornieri