| Authors(s) | Durrell, B. Groombridge, S. Tonge, and Q. Bloxam. 1989. Pyxis arachnoides. Madagascar spider tortoise, Tsakafy, Kapila. In: I. R. Swingland, and M. W. Klemens (eds.). The Conservation Biology of Tortoises, pp. 103-104. Occ. Pap. IUCN/SSC 5, Tortoise and Freshwater Specialists Group. IUCN, Gland, Switzerland. |