| Authors(s) | Hayes, F. E., K. R. Beaman, W. K. Hayes, and L. E. Harris, Jr. |
| Year | 1988 |
| Title | Defensive behavior in the Galapagos tortoise (Geochelone elephantopus), with comments on the evolution of insular gigantism. Herpetologica 44: 11-17. |
| Authors(s) | Hayes, F. E., K. R. Beaman, W. K. Hayes, and L. E. Harris, Jr. |
| Year | 1988 |
| Title | Defensive behavior in the Galapagos tortoise (Geochelone elephantopus), with comments on the evolution of insular gigantism. Herpetologica 44: 11-17. |